Vatanmniuk O K，Mari S，Lu Y P，et al． AtPCS1，a phytochelatin synthase from Arabidopsis:isolation and in vitro reconstitution[J]. Proc Natl Acad Sci，1999，96:7l10－7l15.
[2]
Ha S B，Smith A P，Howden R，et al． Phytochelatin synthase genes from Arabidopsis and the yeast Schizosaccharomyces pombe[J]. Plant Cell，1999，11:l153－ l164.
[3]
Huang H G，Li T X，Yang X E，et al． Research advances in?plant lead tolerance and detoxification mechanism[J]. Ying?Yong Sheng Tai Xue Bao，2009，20(3):696－704.
[4]
?Liu GY，Zhang Y X，Chai TY. Phytochelatin synthase of?Thlaspi caerulescens enhanced tolerance and accumulation of?heavy metals when expressed in yeast and tobacco[J]. Plant?Cell Rep，2011，30(6):1067－1076.
[5]
Osaki Y，Shirabe T，Nakanishi H，et al． Characterization of?phytochelatin synthase produced by the primitive red alga?Cyanidioschyzon merolae[J]. Metallomics，2009，1(4):?353－358.
[6]
?Wojas S，Clemens S，Sk?odowska A，et al． Arsenic response of AtPCS1-and CePCS-expressing plants -effects?of external As (V) concentration on As-accumulation?pattern and NPT metabolism[J]. J Plant Physiol，2010, 167(3):169－175.
[7]
Wojas S，Ruszczyńska A，Bulska E，et al． The role of subcellular distribution of cadmium and phytochelatins in the?generation of distinct phenotypes of AtPCS1-and CePCS3expressing tobacco[J]. J Plant Physiol，2010，167(12):?981－988.
[8]
?Meyer C L，Peisker D，Courbot M，et al． Isolation and?characterization of Arabidopsis halleri and Thlaspi caerulescens phytochelatin synthases[J]. Planta，2011，234?(1):83－95.
[9]
Brunetti P，Zanella L，Proia A，et al． Cadmium tolerance?and phytochelatin content of Arabidopsis seedlings overexpressing the phytochelatin synthase gene AtPCS1[J]. J?Exp Bot，2011，62(15):5509－5519.
[10]
Zhu Y L，Pilon-Smiths E A H，Tarun A S，et al． Overexpression of glutathione synthetase in Indian mustard enhances cadmium accumulation and tolerance[J]. Plant?Physiol，1999，119:73－79.
[11]
Gisbert C，Ros R，De Haro A，et al． A plant genetically?modified that accumulates Pb is especially promising for?phytoremediation[J]. Biochem Biophys Res Commun, 2003，303:440－445.
[12]
范洪黎 . 苋菜超积累镉的生理机制研究[D]. 北京:?中国农业科学院研究生院，2007.
[13]
Zhao F J，Jiang R F，Dunham S J，et al． Cadmium uptake, translocation and tolerance in the hyperaccumulator Arabidopsis halleri[J]. New Phytol，2006，172:646－654.
[14]
?Garbisu C，Alkorta I. Phytoextraction: a cost-effective?plant-based technology for the removal of metals from?the environment[J]. Bioresource Technoi，2001，77?(3):229－236.
[15]
Wang H C，Wu J S，Chia J C，et al． Phytochelatin synthase is regulated by protein phosphorylation at a threonine residue near its catalytic site[J]. J Agric Food?Chem，2009，57(16):7348－7355.
[16]
Vestergaard M，Matsumoto S，Nishikori S，et al． Chelation?of cadmium ions by phytochelatin synthase:role of the?cysteine-rich C-terminal[J]. Anal Sci，2008，24(2):277?－281.
