Continuum of Gene-Expression Profiles Provides Spatial Division of Labor within a Differentiated Cell Type

？ Multi-tasking theory explains continuum gene-expression within a cell type ？ Enterocytes show a 1D expression continuum and trade-off between three tasks ？ Hepatocytes show a 3D continuum and four complexes of liver tasks ？ Spatial zonation of task-specialist cells arises from task-performance gradient