32 Hou X G, Ramsk？ld L, Bergstr？m J. Composition and preservation of the Chengjiang fauna—A Lower Cambrian soft-bodied biota. Zool Scr, 1991, 20: 395—411
[3]
33 Liu J N, Shu D G, Han J, et al. The lobopod Onychodictyon from the Lower Cambrian Lagerst？tte revisited. Acta Palaeontol Pol, 2008, 53: 285—292
[4]
34 Hou X G, Bergstr？m J. Cambrian lobopodians-ancestors of extant onychophorans? Zool J Linn Soc, 1995, 114: 3—19
[5]
35 Chen J Y, Zhou G Q, Ramsk？ld L. A new Early Cambrian Onychophoran-like animal, Paucipodia gen. nov., from the Chengjiang fauna, China. Trans R Soc Edinb-Earth Sci, 1995, 85: 275—282
[6]
36 Hou X G, Ma X Y, Zhao J, et al. The lobopodian Paucipodia inermis from the Lower Cambrian Chengjiang fauna, Yunnan, China. Lethaia, 2004, 37: 235—244
[7]
37 Liu J N, Shu D G, Han J, et al. Morpho-anatomy of the lobopod Megadictyon cf. haikouensis from the Early Cambrian Chengjiang Lagerst？tte, South China. Acta Zool, 2007, 88: 279—288
[8]
38 Liu J N, Shu D G, Han J, et al. A large Xenusiid lobopod with complex appendages from the Lower Cambrian Chengjiang Lagerst？tte. Acta Palaeontol Pol, 2006, 51: 215—222
[9]
39 Dzik J, Krumbiegel G. The oldest ‘onychophoran’ xenusion: A link connecting phyla? Lethaia, 1989, 22: 169—181
[10]
40 Bergstr？m J, Hou X G. Cambrian onychophora or xenusians. Zool Anz, 2001, 240: 237—245
[11]
41 Budd G. Tardigrades as ‘Stem-Group Arthropods’: The evidence from the Cambrian Fauna. Zool Anz, 2001, 240: 265—279
[12]
1 Collins D. Misadventures in the Burgess shale. Nature, 2009, 460: 952—953
5 Hou X G, Aldridge R J, Bergstr？m J, et al. The Cambrian Fossils of Chengjiang, China: The Flowering of Early Animal Life. Oxford: Blackwell Publishing, 2004. 233
9 Han J, Zhang Z F, Shu D G. Discovery of the proboscis on Tylotites petiolaris. Northwestern Geol, 2003, 36: 87—93
[21]
10 Huang D Y, Vannier J, Chen J Y. Anatomy and lifestyles of Early Cambrian priapulid worms exemplified by Corynetis and Anningvermis from the Maotianshan Shale (SW China). Lethaia, 2004, 37: 21—33
13 Huang D Y, Vannier J, Chen J Y. Recent Priapulidae and their Early Cambrian ancestors: Comparisons and evolutionary significance. Geobios, 2004, 37: 217—228
[25]
14 Han J, Zhang X L, Zhang Z F, et al. A new platy-armored worm from the Early Cambrian Chengjiang Lagerst？tte, South China. Acta Geol Sin-Engl Ed, 2003, 77: 1—6
16 Hou X G, Berstr？m J, Yang J. Distinguishing anomalocaridids from arthropods and priapulids. Geol J, 2006, 41: 259—269
[28]
17 Han J, Liu J N, Zhang Z F, et al. Trunk ornament on the palaeoscolecid worms Cricocosmia and Tabelliscolex from the Early Cambrian Chengjiang deposits of China. Acta Palaeontol Pol, 2007, 52: 423—431
[29]
18 Huang D Y. Early Cambrian worms from SW China: Morphology, systematics, lifestyles and evolutionary significance. Doctor Dissertation. Lyon: Université Claude Bernard Lyon, 2005
[30]
19 Ma X Y, Hou X G, Siveter D J, et al. A new priapulid-like worm from the Early Cambrian Chengjiang Lagerst？tte, China. In: Abstracts of the 51st Annual Meeting of the Palaeontological Association, 2007 Dec 16—19, Uppsala. Palaeontol Assoc Newsl, 2007, 66: 43
[31]
20 Maas A, Huang D Y, Chen J Y, et al. Maotianshan-Shale nemathelminths: Morphology, biology, and the phylogeny of Nemathelminthes. Palaeogeogr Palaeoclimatol Palaeoecol, 2007, 254: 288—306
[32]
21 Whittard W F. Palaeoscolex piscatorum gen. et sp. nov., a worm from the Tremadocian of Shropshire. Q J Geol Soc Lond, 1953, 109: 125—136
[33]
22 Conway M S. The cuticular structure of a 495-Myr-old type species of the fossil worm Palaeoscolex, P. piscatorum (?Priapulida). Zool J Linn Soc, 1997, 119: 69—82
[34]
23 Chen J Y, Zhou G Q. Biology of the Chengjiang Fauna. Bull Natl Mus Nat Sci Taiwan, 1997, 10: 11—106
[35]
24 Harvey T H P, Dong X P, Donoghue C J. Are palaeoscolecids ancestral ecdysozoans? Evol Dev, 2010, 12: 177—200
[36]
25 黄迪颖, 陈均远, Vannier J. 早寒武世曳鳃形目化石分类问题的讨论. 科学通报, 2006, 51: 243—249
27 Dong X P, Donoghue P C J, Cunningham J A, et al. The anatomy, affinity, and phylogenetic significance of Markuelia. Evol Dev, 2005, 7: 468—482
[39]
28 侯先光, 陈均远. 云南澄江早寒武世节肢动物与环节中间性生物——Luolishania gen. nov. 古生物学报, 1989, 28: 207—213
[40]
29 Ma X Y, Hou X G, Bergstr？m J. Morphology of Luolishania longicruris (Lower Cambrian, Chengjiang Lagerst？tte, SW China) and the phylogenetic relationships within lobopodians. Arthropod Struct Dev, 2009, 38: 271—291
42 Huang D Y, Chen J Y, Vannier J, et al. Early Cambrian sipunculan worms from southwest China. Proc R Soc B-Biol Sci, 2004, 271: 1671—1676
[43]
43 Chen J Y, Huang D Y. A possible Lower Cambrian Cheatognath (arrow worm). Science, 2002, 298: 187
[44]
44 Hu S X. Taphonomy and palaeoecology of the Early Cambrian Chengjiang biota from eastern Yunnan, China. Berliner Pal？obiologische Abhandlungen, 2005, 7: 1—197
46 Chapman G. The hydrostatic skeleton in the invertebrates. Biol Rev, 1958, 33: 338—371
[47]
47 Dornbos S Q, Chen J Y. Community palaeoecology of the early Cambrian Maotianshan Shale biota: Ecological dominance of priapulids worms. Palaeogeogr Palaeoclimatol Palaeoecol, 2008, 258: 200—212
[48]
48 Zhang X G, Hou X G, Bergstr？m J. Early Cambrian priapulid worms buried with their lined burrows. Geol Mag, 2006, 143: 743—748
[49]
49 Hammond R A. The burrowing of Priapulus caudatus. J Zool Lond, 1970, 162: 469—480
[50]
50 Ruppert E E, Barnes R D. Invertebrate Zoology. 6th ed. New York: Sanders College Publishing, 1996. 1100
[51]
51 Han J, Zhang Z F, Liu J N. Evidence of priapulid scavenging from the Early Cambrian Chengjiang deposits, Southern China. Palaios, 2007, 22: 691—694 1134
