Taxonomy of new relatives of Onthophagus catenatus Lansberge, 1883 from New Guinea (Coleoptera, Scarabaeidae, Scarabaeinae)

Four new taxa from New Guinea are proposed in the dung beetle genus Onthophagus Latreille, 1802, all in the operational group of O. catenatus Lansberge, 1883. The group is discussed, defined, and the five taxa included are listed, keyed, and diagnosed. Three new species are described: Onthophagus abmisibilus (from West New Guinea, Indonesia), O. kokodanus, O. kokosquamatus (both from Papua New Guinea). One new species comprises a lowland and an upland subspecies: O. kokodanus kokodanus and kokodanus hagenaltus (both in Papua New Guinea).