| [1] | Darwin C (1871) The descent of man and selection in relation to sex. London: Murray.
|
| [2] | Bateman AJ (1948) Intrasexual selection in Drosophila. Heredity 2: 349–368.
|
| [3] | Trivers R (1972) Parental investment and sexual selection. In: Campbell BG, editor. Sexual selection and the descent of man: the Darwinian pivot. New Brunswick, New Jersey: Transaction Publishers. pp. 136–179.
|
| [4] | Andersson M (1994) Sexual selection. Princeton, NJ: Princeton University Press.
|
| [5] | Shuster SM, Wade MJ (2003) Mating systems and strategies. Princeton Univ Press.
|
| [6] | Berglund A, Bisazza A, Pilastro A (1996) Armaments and ornaments: an evolutionary explanation of traits of dual Utility. Biological Journal of the Linnean Society 58: 385–399.
|
| [7] | Tomkins JL, Simmons LW (1998) Female choice and manipulations of forceps size and symmetry in the earwigForficula auriculariaL. Animal Behaviour 56: 347–356.
|
| [8] | Pope DS (2000) Testing function of fiddler crab claw waving by manipulating social context. Behavioral Ecology and Sociobiology 47: 432–437.
|
| [9] | Andersson M, Simmons L (2006) Sexual selection and mate choice. Trends in Ecology & Evolution 21: 296–302.
|
| [10] | Emlen D (2008) The evolution of animal weapons. Annual Review of Ecology, Evolution and Systematics 39: 387–413.
|
| [11] | Zahavi A (1975) Mate Selection - Selection for a Handicap. Journal of Theoretical Biology 53: 205–214.
|
| [12] | Grafen A (1990) Sexual selection unhandicapped by the Fisher process. Journal of Theoretical Biology 144: 473–516.
|
| [13] | Bonduriansky R, Day T (2003) The evolution of static allometry in sexually selected traits. Evolution 57: 2450–2458.
|
| [14] | Nur N, Hasson O (1984) Phenotypic plasticity and the handicap principle. Journal of Theoretical Biology 110: 275–297.
|
| [15] | Kodric-Brown A, Sibly RM, Brown JH (2006) The allometry of ornaments and weapons. Proceedings of the National Academy of Sciences of the United States of America 103: 8733–8738.
|
| [16] | Hamilton W, Zuk M (1982) Heritable true fitness and bright birds: a role for parasites? Science 218: 384.
|
| [17] | Moller A, Christe P, Lux E (1999) Parasitism, host immune function, and sexual selection. Quarterly Review of Biology 74: 3–20.
|
| [18] | Moller AP (1990) Parasites and sexual selection - current status of the Hamilton and Zuk hypothesis. Journal of Evolutionary Biology 3: 319–328.
|
| [19] | Howard RD, Minchella DJ (1990) Parasitism and Mate Competition. Oikos 58: 120–122.
|
| [20] | Pomfret J, Knell R (2006) Immunity and the expression of a secondary sexual trait in a horned beetle. Behavioral Ecology 17: 466.
|
| [21] | Rantala M, Roff D, Rantala L (2007) Forceps size and immune function in the earwig Forficula auricularia L. Biological Journal of the Linnean Society 90: 509–516.
|
| [22] | Ezenwa VO, Jolles AE (2008) Horns honestly advertise parasite infection in male and female African buffalo. Animal Behaviour 75: 2013–2021.
|
| [23] | Rantala M, Koskim？ki J, Taskinen J, Tynkkynen K, Suhonen J (2000) Immunocompetence, developmental stability and wingspot size in the damselfly Calopteryx splendens L. Proceedings of the Royal Society of London Series B: Biological Sciences 267: 2453.
|
| [24] | Siva-Jothy M (2000) A mechanistic link between parasite resistance and expression of a sexually selected trait in a damselfly. Proceedings of the Royal Society of London Series B: Biological Sciences 267: 2523.
|
| [25] | Contreras-Garduno J, Lanz-Mendoza H, Cordoba-Aguilar A (2007) The expression of a sexually selected trait correlates with different immune defense components and survival in males of the American rubyspot. Journal of insect physiology 53: 612–621.
|
| [26] | Rantala M, Kortet R (2003) Courtship song and immune function in the field cricket Gryllus bimaculatus. Biological Journal of the Linnean Society 79: 503–510.
|
| [27] | Ryder J, Siva-Jothy M (2000) Male calling song provides a reliable signal of immune function in a cricket. Proceedings of the Royal Society of London Series B: Biological Sciences 267: 1171.
|
| [28] | Simmons L, Zuk M, Rotenberry J (2005) Immune function reflected in calling song characteristics in a natural population of the cricket Teleogryllus commodus. Animal Behaviour 69: 1235–1241.
|
| [29] | Rantala M, Jokinen I, Kortet R, Vainikka A, Suhonen J (2002) Do pheromones reveal male immunocompetence? Proceedings of the Royal Society of London Series B: Biological Sciences 269: 1681.
|
| [30] | Cotter S, Kruuk L, Wilson K (2004) Costs of resistance: genetic correlations and potential trade-offs in an insect immune system. Journal of Evolutionary Biology 17: 421–429.
|
| [31] | Latham ADM, Poulin R (2001) Effect of acanthocephalan parasites on the behaviour and coloration of the mud crab Macrophthalmus hirtipes (Brachyura : Ocypodidae). Marine Biology 139: 1147–1154.
|
| [32] | Bollache L, Gambade G, Cezilly F (2001) The effects of two acanthocephalan parasites, Pomphorhynchus laevis and Polymorphus minutus, on pairing success in male Gammarus pulex (Crustacea : Amphipoda). Behavioral Ecology and Sociobiology 49: 296–303.
|
| [33] | Moczek AP (2006) Pupal Remodeling and the Development and Evolution of Sexual Dimorphism in Horned Beetles. American Naturalist 168: 711–729.
|
| [34] | Emlen D (2001) Costs and the diversification of exaggerated animal structures. Science 291: 1534.
|
| [35] | Moczek A, Nijhout H (2004) Trade offs during the Development of Primary and Secondary Sexual Traits in a Horned Beetle. American Naturalist 163: 184–191.
|
| [36] | Simmons L, Emlen D (2006) Evolutionary trade-off between weapons and testes. Proceedings of the National Academy of Sciences, USA 103: 16346.
|
| [37] | Knell R, Pomfret J, Tomkins J (2004) The limits of elaboration: curved allometries reveal the constraints on mandible size in stag beetles. Proceedings of the Royal Society of London Series B: Biological Sciences 271: 523.
|
| [38] | Okada K, Miyatake T (2009) Genetic correlations between weapons, body shape and fighting behaviour in the horned beetle Gnatocerus cornutus. Animal Behaviour 77: 1057–1065.
|
| [39] | Okada K, Miyanoshita A, Miyatake T (2006) Intra-sexual dimorphism in male mandibles and male aggressive behavior in the broad-horned flour beetle Gnatocerus cornutus (Coleoptera: Tenebrionidae). Journal of Insect Behavior 19: 457–467.
|
| [40] | Okada K, Miyatake T (2010) Effect of losing on male fights of broad-horned flour beetle, Gnatocerus cornutus. Behavioral Ecology and Sociobiology 64: 361–369.
|
| [41] | Savvidou N, Bell CH (1994) The effect of larval density, photoperiod and food change on the development of Gnatocerus cornutus (F.)(Coleoptera: Tenebrionidae). Journal of Stored Products Research 30: 17–21.
|
| [42] | Keymer AE (1980) The influence of Hymenolepis diminuta on the survival and fecundity of the intermediate host, Tribolium confusum. Parasitology 81: 405–421.
|
| [43] | Blankespoor C, Pappas P, Eisner T (1997) Impairment of the chemical defence of the beetle, Tenebrio molitor, by metacestodes (cysticercoids) of the tapeworm, Hymenolepis diminuta. Parasitology 115: 105–110.
|
| [44] | Nappi AJ, Christensen BM (2005) Melanogenesis and associated cytotoxic reactions: Applications to insect innate immunity. Insect Biochemistry and Molecular Biology 35: 443–459.
|
| [45] | Mullen LM, Goldsworthy GJ (2006) Immune responses of locusts to challenge with the pathogenic fungus Metarhizium or high doses of laminarin. Journal of insect physiology 52: 389–398.
|
| [46] | Schwarzenbach GA, Ward PI (2007) Phenoloxidase activity and pathogen resistance in yellow dung flies Scathophaga stercoraria. Journal of Evolutionary Biology 20: 2192–2199.
|
| [47] | Nigam Y, Maudlin I, Welburn S, Ratcliffe NA (1997) Detection of phenoloxidase activity in the hemolymph of tsetse flies, refractory and susceptible to infection with Trypanosoma brucei rhodesiense. Journal of Invertebrate Pathology 69: 279–281.
|
| [48] | Hung S, Boucias D (1996) Phenoloxidase Activity in Hemolymph of Naive andBeauveria bassiana-InfectedSpodoptera exiguaLarvae. Journal of Invertebrate Pathology 67: 35–40.
|
| [49] | Newton K, Peters R, Raftos D (2004) Phenoloxidase and QX disease resistance in Sydney rock oysters (Saccostrea glomerata). Developmental and Comparative Immunology 28: 565–569.
|
| [50] | Knell RJ (2009) On the analysis of non-linear allometries. Ecological Entomology 34: 1–11.
|
| [51] | Muggeo VMR (2003) Estimating regression models with unknown break points. Statistics in Medicine 22: 3055–3071.
|
| [52] | Lawniczak MKN, Barnes AI, Linklater JR, Boone JM, Wigby S, et al. (2007) Mating and immunity in invertebrates. Trends in Ecology & Evolution 22: 48–55.
|
| [53] | Folstad I, Karter AJ (1992) Parasites, bright males, and the immunocompetence handicap. American Naturalist 139: 603–622.
|
| [54] | Yamane T, Okada K, Nakayama S, Miyatake T (2010) Dispersal and ejaculatory strategies associated with exaggeration of weapon in an armed beetle. Proceedings of the Royal Society of London Series B: Biological Sciences.
|
| [55] | Harano T, Okada K, Nakayama S, Miyatake T, Hosken DJ (2010) Intralocus Sexual Conflict Unresolved by Sex-Limited Trait Expression. Current Biology 20: 2036–2039.
|
| [56] | Okada K, Yamane T, Miyatake T (2010) Ejaculatory strategies associated with experience of losing. Biology Letters 6: 593.
|
| [57] | Hurd H, Arme C (1986) Hymenolepis diminuta: effect of metacestodes on production and viability of eggs in the intermediate host, Tenebrio molitor. Journal of Invertebrate Pathology 47: 225–230.
|
| [58] | Hurd H, Ardin R (2003) Infection increases the value of nuptial gifts, and hence male reproductive success, in the Hymenolepis diminuta-Tenebrio molitor association. Proceedings of the Royal Society of London Series B: Biological Sciences 270: S172–S174.
|
| [59] | Hurd H, Fogo S (1991) Changes induced by Hymenolepis diminuta (Cestoda) in the behaviour of the intermediate host Tenebrio molitor (Coleoptera). Canadian Journal of Zoology 69: 2291–2294.
|
| [60] | Hurd H, Warr E, Polwart A (2001) A parasite that increases host lifespan. Proceedings of the Royal Society of London Series B: Biological Sciences 268: 1749.
|
| [61] | Sheldon BC, Verhulst S (1996) Ecological immunology: Costly parasite defences and trade-offs in evolutionary ecology. Trends in Ecology & Evolution 11: 317–321.
|
| [62] | Zuk M, Stoehr AM (2002) Immune Defense and Host Life History. American Naturalist 160: S9–S22.
|
| [63] | Rantala MJ, Vainikka A, Kortet R (2003) The role of juvenile hormone in immune function and pheromone production trade-offs: a test of the immunocompetence handicap principle. Proceedings of the Royal Society of London Series B: Biological Sciences 270: 2257–2261.
|
| [64] | von Schantz T, Bensch S, Grahn M, Hasselquist D, Wittzell H (1999) Good genes, oxidative stress and condition-dependent sexual signals. Proceedings of the Royal Society of London Series B: Biological Sciences 266: 1.
|
| [65] | Dowling D, Simmons L (2009) Reactive oxygen species as universal constraints in life-history evolution. Proceedings of the Royal Society B.
|
| [66] | Schmid-Hempel P (2005) Evolutionary ecology of insect immune defenses. Annual Review of Entomology 50: 529–551.
|
| [67] | Cole TJ, Eggleston P, Hurd H (2003) Juvenile hormone titre and egg production in Tenebrio molitor infected by Hymenolepis diminuta: effect of male and/or female infection, male age and mating. Journal of Insect Physiology 49: 583–590.
|
